B.S., Molecular Biophysics & Biochemistry - Yale University / Yale University, 1987
M.D. - Stanford University School of Medicine, 1995
Ph.D. (Biochemistry) - Stanford University School of Medicine, 1995
Postdoc (Aging Biology) - Massachusetts Institute of Technology, 2001
Residency in Clinical Pathology - Brigham and Women's Hospital, 2001
Medical degree, DO - Stanford University School of Medicine, 1995